| Map distance from the gene to centromere = | # of MII asci/2 | x 100 |
| total # of asci |
| Linked genes: | PD asci= NPD asci |
| NPD asci / Tetratype asci > 1/4 |
| Unlinked genes: | PD asci >> NPD asci |
| NPD asci / Tetratype asci <1/4 |
| Map distance between 2 genes: | (# of Tetratype asci / 2) + (# of NPD asci) | x 100 |
| total # of asci |